Education and Employment
| September 1980 - April 1984 | B.Sc. (Hons.), Life Sciences, Queen's University, Kingston, Ontario |
| July 1984 - April 1987 | M.Sc., Medical Physiology, University of Calgary, Calgary, Alberta |
| May 1987 - December 1990 | Ph.D. program, Physiology (Pharmacology Track) Southern Illinois University, Springfield, Illinois |
| January 1990 - December 1992 | Ph.D., Biomedical Sciences (Physiology) Creighton University and Boys Town National Research Hospital, Omaha, Nebraska |
| June 1993 - December 1995 | Postdoctoral Fellow, Department of Otolaryngology, Johns Hopkins University School of Medicine, Baltimore, Maryland |
| January 1996 – August 1997 | Postdoctoral Fellow, Department of Anatomical Sciences and Neurobiology, University of Louisville School of Medicine, Louisville, Kentucky |
| September 1997 – August 2003 | Assistant Professor, Departments of Pharmacology and Medical and Molecular Physiology, University of Minnesota School of Medicine, Duluth, Minnesota |
| August 2003-present | Associate Professor (with tenure), Department of Pharmacology and Physiology, University of Minnesota Medical School, Duluth, Minnesota |
Professional Memberships
Society for Neuroscience 1985-present
Association for Research in Otolaryngology 1992-present
Sigma Xi 1998-present
American Physiological Society 2002-present
American Society for Pharmacology and Experimental Therapeutics 2002-present